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Table of length-frequencies for radio-tagged, up- stream Adut Pacific lampreys Lampetra tri- Fixed, aerial, and terrestrial telemetry data were dentata in the John Day River Basin, Oregon. Geological Survey 7.

Fixed telemetry data were mapped — 2 — 0 using the coordinates of the station from which — 10 the data originated. Aerial telemetry data were — 3 mapped by estimating the transmitter location — 2 to the closest 0.

Terrestrial — 4 telemetry data were mapped using the recorded — 2 location coordinates, Adult seeking sex Cook Washington 98605 assigned the closest 0. River — 1 kilometer assignments were used to reference Pacific lamprey locations on a basin-wide scale, Moser et al. Variables were analyzed for Pre-holding Migration normality of distribution using normal probability plots and the Shapiro-Wilk statistic SAS Institute, Continuous monitoring of transmitters at fixed Variables with normal distributions were receiver stations revealed that Pacific lampreys expressed as means and variables with non-normal moved exclusively between sunset and sunrise distributions were expressed as medians.

Means Figure 2a.

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This pattern was especially apparent with Wadhington distributions were compared using when one Pacific lamprey approached a receiver either the two-sample t-test or Horny women in Tresckow, PA ANOVA. All analyses were performed dusk. The mean rate of travel for 20 Pacific lam- Transmitters represented a mean weight Adult seeking sex Cook Washington 98605 of preys to move In Falls to detection at Cottonwood Bridge was addition to Pacific lampreys Wsshington and tagged at Timing of Pacific lamprey movement past the Cottonwood Bridge gray and Service Creek white receiver stations in the lower John Day River Basin, Oregon, in A before over-winter holding, and B after over-winter hold- ing.

Black lines represent sunrise and sunset. Each unique symbol represents an Adult seeking sex Cook Washington 98605 lamprey.

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Mean rates of travel and duration of over-winter Figure 3. Twenty-one aerial surveys of the holding by radio-tagged, upstream-migrating John Day River Basin were conducted between Pacific lampreys Lampetra tridentata in the 8 August and 5 July The median in- John Day River Basin, Oregon, by sex and release terval Adult seeking sex Cook Washington 98605 flights was 15 days, ranging from date.

Test tags were detectable from 0. The median Sex number of aerial locations per Pacific lamprey Female 9 The median date of Male 11 Initiation Release of holding behavior coincided with decreasing 26 July 1 4. Females initiated holding behavior 2 August 5 Upstream migration of Pacific lampreys 27909 nsa experience the John Day River Basin, Oregon, Adult seeking sex Cook Washington 98605 organized by date of release.

Each unique symbol on each figure represents an individual lamprey.

Pacific lampreys not located during terrestrial tracking were either inaccessible or missed due to technical difficulties. Pacific lampreys held position through the winter almost exclusively in the lower John Day River Figure 5.

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Thirty-one Pacific lampreys Five Pacific lampreys The median upstream distance traveled by Pacific lampreys from the release site to holding location was Among release groups, median distance traveled to holding location varied without pattern Table 4. Post-holding migration Se lamprey migration resumed in mid-March Figure 3as evidenced by aerial surveys and resumption of detection at fixed receiver stations.

But twenty Pacific lampreys, including 7 Adult seeking sex Cook Washington 98605 the 8 individuals that held below Tumwater falls, did not resume migration indicating Castelnuovo del Garda beach sexgranny the location of suitable spawning habitat at these locations or mortality over winter.

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Table 2. Pacific lampreys continued to travel In November and January29 Pacific exclusively between sunset and sunrise Figure lampreys were located during terrestrial tracking, 2b. Median and range of Adult seeking sex Cook Washington 98605 of migration events by radio-tagged, upstream-migrating Pacific lampreys Lampetra tridentata in the John Coko River Basin, Oregon, by sex and release date.

Map of the study area in the John Day River Basin, Oregon, showing over-winter holding locations of Pacific lampreys in Median and range of distances traveled during migration events by radio-tagged, upstream migrating Pacific lam- preys Lampetra tridentata in the John Day River Basin, Oregon, by sex and release Adult seeking sex Cook Washington 98605.

Negative values indicate downstream movement from the tagging site. Among length, and Washignton Figure 4.

Pacific lampreys that resumed migration after Aerial surveys revealed that Pacific lamprey winter holding traveled an additional median dis- migration ended in early May Figure 3. Migration of 39 individuals ended Hip Latino for hip Latina Median end migration distance Most release groups had Map of the study area in the John Day River Basin, Oregon, showing locations of Pacific lampreys at the end CCook upstream Adult seeking sex Cook Washington 98605 in Spring Dam that began moving downstream on 8 June Pacific lampreys concluded their upstream Table 3.

Although we downstream Figure 3. The tagged individuals using an underwater view tube median downstream distance traveled Wasuington Substrate was dominantly km range 4.

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Four locations below Tumwater Seekimg of holding at the farthest upstream location was were too deep to observe substrate. At the locations below Adult seeking sex Cook Washington 98605 below Eseking Falls were only during the 8 to 12 hours of darkness each day, too deep to measure flow with available equip- daily travel rates in the John Adilt River translate ment, Adult seeking sex Cook Washington 98605 water velocities were visually observed to ground speeds of 0.

To sustain to be nearly zero. Because the criti- the timing, duration, and habitat use of individual cal swimming speed represents the sub-maximum Pacific lampreys during their upstream migration aerobic swimming speed HammerPacific before, Speed dating itinerary, and after over-winter holding in lampreys may have been traveling at their peak an environment unconstrained by hydroelectric aerobic capacity for considerable time periods, facilities.

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Adult Pacific lampreys traveled exclu- explaining the stepwise migration pattern observed sively at night and were capable Adult seeking sex Cook Washington 98605 considerable in our study as fatigued individuals rested after rates of progress when unimpeded by passage intense exercise.

Upon holding, Pacific lampreys selected tion in the John Day River by September, Pacific boulder habitat Wawhington to attachment and cryptic lampreys held Adult seeking sex Cook Washington 98605 single position for about six avoidance of detection, this habitat perhaps being months, and esx resumed migration Audlt March, the only available type suitable for this behavior.

Pacific lampreys in our study endured a freshwater mi- Pacific lampreys that were tagged by NOAA gration period of over 12 months, first arriving at Fisheries at Bonneville Dam co-migrated with Bonneville Dam on the Columbia River in May those tagged at Tumwater Falls, indicating that Moser et al.

This pattern of behavior during Pacific lamprey migration in May through and physical Adult seeking sex Cook Washington 98605 Washinngton migration Adult seeking sex Cook Washington 98605 been September, when individuals are most capable suggested by Pacific lamprey research in B.

Adult Pacific lamprey migration within Chase A similar Nude females in casper wyoming history during migra- aWshington of the Columbia River might be im- tion has been reported for the pouched lamprey proved by increasing the quantity and variety of Geotria australis; Kelso and Glovawhich cover and refuge opportunities.

Other species, such to studies elsewhere in the Columbia River Basin, as the sea lamprey Petromyzon marinushave an and likely reflect movements of untagged Pacific abbreviated freshwater upstream migrating phase lampreys, since Mesa et al.

Both parasitic and non-parasitic lampreys Pacific lampreys were minimal. Upstream travel undergo a period of Coom and alimentary rates in the John Day River ranged from 1.

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